Loach catfish

Doumea cf typica from the Dja in southern Cameroon.

The Quappenwelse ( Amphiliidae ) are a family of fish from the order of catfish -like ( Siluriformes ). They are widespread in tropical Africa and are found in fast-flowing waters of higher layers most frequently.

  • 3.1 Notes and references


Quappenwelse are small fish of mostly less than 12 and up to 19 centimeters in length. Your body is elongated with a pointed head and a slender, long caudal peduncle. The animals have three pairs of barbels, the nasal pair is missing. The pectoral and ventral fins are wide in many species, forming a weak suction cup with which the animals can cling to rocks. The dorsal and anal fins have a short base. Hard jets is missing in all fins, except for Trachyglanis and Leptoglaninae where breast and dorsal fin have a poorly developed hard radiation. A long, flat grease fin is present in some species and has a modified scale, which serves as a rigid beam. The swim bladder is regressed.

There are three subfamilies:

  • The Amphiliinae have a relatively short body with inferior mouth. They have never bone plates or a neck shield.
  • The Doumeinae have an elongated body with inferior mouth, which has a neck shield and often bone plates.
  • The Leptoglaninae the maxilla is greatly extended, and in contrast to most catfish, the proximal radials are connected.


The monophyly of the Quappenwelse is doubted by some authors. This then arrange the Doumeinae and Leptoglaninae into their own family Doumenidae. In the classical system, the Quappenwelse be placed in the superfamily Loricarioidea. After molecular biological studies, they include within the subordination of Siluroidei for "Big Africa " group.

Without spin-off of Doumeinae and Leptoglaninae the family are attributed to 12 genera with 66 species.

Subfamily Amphiliinae

The Quappenwelse the subfamily Amphiliinae have relatively short body, similar to those of Plattschmerlen ( Balitoridae ). The mouth is inferior, bone plates and a neck shield missing.

  • Genus Amphilius Amphilius atesuensis
  • Amphilius Baudoni
  • Amphilius brevis
  • Amphilius cryptobullatus
  • Amphilius jacksonii
  • Amphilius kakrimensis
  • Amphilius kivuensis
  • Amphilius Lamanites
  • Amphilius lampei
  • Amphilius laticaudatus
  • Amphilius lentiginosus
  • Amphilius longirostris
  • Amphilius maesii
  • Amphilius natalensis
  • Amphilius opisthophthalmus
  • Amphilius platychir
  • Amphilius pulcher
  • Amphilius rheophilus
  • Amphilius uranoscopus
  • Amphilius zairensis
  • Paramphilius firestonei
  • Paramphilius Goodi
  • Paramphilius teugelsi
  • Paramphilius trichomycteroides

Subfamily Doumeinae

The Quappenwelse the subfamily Doumeinae have an exceptionally long maxilla. The fin located towards the body carriers are fused together.

  • Genus Andersonia Andersonia leptura
  • Belonoglanis brieni
  • Belonoglanis tenuis
  • Doumea alula
  • Doumea angolensis
  • Doumea chappuisi
  • Doumea skeltoni
  • Doumea thysi
  • Doumea typica
  • Phractura ansorgii
  • Phractura bovei
  • Phractura brevicauda
  • Phractura clauseni
  • Phractura fasciata
  • Phractura gladysae
  • Phractura intermedia
  • Phractura lindica
  • Phractura longicauda
  • Phractura macrura
  • Phractura scaphyrhynchura
  • Phractura tenuicauda
  • Trachyglanis ineac
  • Trachyglanis intermedius
  • Trachyglanis minutus
  • Trachyglanis sanghensis

Subfamily Leptoglaninae

The Quappenwelse the subfamily Leptoglaninae have elongated body. The mouth is inferior. The neck is protected by a bony plate. Also along the trunk, often bone plates.

  • Genus Dolichamphilius Dolichamphilius brieni
  • Dolichamphilius longiceps
  • Leptoglanis bouilloni
  • Leptoglanis camerunensis
  • Leptoglanis Dorae
  • Leptoglanis flavomaculatus
  • Leptoglanis mandevillei
  • Leptoglanis rotundiceps
  • Leptoglanis xenognathus
  • Psammphiletria delicata
  • Psammphiletria nasuta
  • Tetracamphilius angustifrons
  • Tetracamphilius clandestinus
  • Tetracamphilius notatus
  • Tetracamphilius pectinatus
  • Zaireichthys heterurus
  • Zaireichthys zonatus


  • Joseph S. Nelson: Fishes of the world. 4 edition. John Wiley & Sons, Hoboken 2006, ISBN 978-0-471-25031-9, p.164
  • Tim M. Berra: Freshwater Fish Distribution. The University of Chicago Press, Chicago 2007, ISBN 9780226044422, pp. 216-217.